Publications

For a full list of papers and citations see: H.M. Kharouba's Google Scholar profile.
Underlined authors are students or postdocs in the lab.

2020 - 2025

Yannelli, F. +20 co-authors (incl. Kharouba, H.M.). 2025. Ecology for a social revolution: Re-defining the role of ecological and environmental science professionals and their responsibilities towards society. Research Ideas and Outcomes 11: e152859. link

Panchen, Z.A., Doubt, J., Saarela, J.M., Kharouba, H.M. 2025. Digitised herbarium specimen data reveal a climate change-related trend to an earlier, shorter Canadian Arctic flowering season, and phylogenetic signal in Arctic flowering times. New Phytologist. link

Pekos, Z.Rivest, S.A., Mitchell, G., and Kharouba, H.M. 2025. Seasonality of native and non-native flowers does not influence butterfly nectar foraging decisions in a semi-urban meadow habitat. Insect Conservation and Diversity.  link

Ortiz, A.M.D. +23 co-authors (incl. Kharouba, H.M.). 2025. Listening deeply to indigenous people: A collaborative perspective and reflection between a Mapuche Machi and ecologists. Ecology and Evolution. link

Dolson, S. and Kharouba, H.M. 2024. 30 years of terrestrial insect richness patterns across elevation: What have we learned? A global meta-analysis. Journal of Animal Ecology. pdf

Loughnan, D., Joly, S., Legault, G., Kharouba, H. M., Betancourt, M., & Wolkovich, E. M. 2024. Phenology varies with phylogeny but not by trophic level with climate change. Nature Ecology & Evolution, 1-8. pdf

Keefe, H. and Kharouba, H.M. 2024. Growing degree-days do not explain moth species distributions at broad scales. Ecosphere. link

Kharouba, H.M. 2024. Shifting the paradigm: the role of introduced plants in the resiliency of terrestrial ecosystems to climate change. Global Change Biology. link

Kharouba, H.M. and Williams, J.L. 2024. Forecasting species' responses to climate change using space-for-time substitution. Trends in Ecology & Evolutionlink  pdf

Kharouba, H.M. 2024. (Editorial) Now is the time for academics to think and act beyond academia. FACETS.  link ​

Rivest, S.A., and Kharouba, H.M. 2024. Taxonomic and functional homogenization of butterfly communities along an urban gradient. Insect Conservation and Diversity. link

Kharouba, H.M., and Wolkovich, E.M. 2023. Lack of evidence for the match-mismatch hypothesis across terrestrial trophic interactions. Ecology Letters.  link

Robinson, M.L. +191 co-authors (incl. Kharouba, H.M.)  2023. Plant size, latitude, and phylogeny explain within-population variability in herbivory. Science. link

Reich, M.S., Kindra, M.K., Dargent, F., Hu, L., Flockhart, D.T.T, Norris, D.R., Kharouba, H.M., Talavera, G., and Bataille, C.P. 2023Metals and metal isotopes in insect wings: Implications for diet, geolocation and pollution exposure. Frontiers in Ecology and Evolution.  link  press release     

Rivest, S.A., Wolkovich, E.M., Kharouba, H.M. 2023. Flowering phenology influences butterfly visitation to non-native plants for nectar in an oak savanna. Ecology.  link 

A graphic with various species of butterflies and flowers demonstrating that flowering phenology influences butterfly visitation to non-native plants for nectar in an oak savanna.

Foster, S., Kharouba, H.M., and Smith, T.W. 2022. Testing the assumption of environmental equilibrium in an invasive plant species over a 130 year history. Ecography. link graphical abstract cover plain language summary

Gilmour, S. and Kharouba, H.M. 2022. Eastern monarch larval performance may not be affected by shifts in phenological synchrony with milkweed. Ecology and Evolution. link  graphical abstract

Kharouba, H.M. and Yang, L.H. 2021. Disentangling the direct, indirect and combined effects of experimental warming on a plant-insect herbivore interaction. Ecosphere 12:e03778.  link

Rivest, S.A. and Kharouba, H.M. 2021Anthropogenic disturbance promotes the abundance of a newly introduced butterfly in Canada, Polyommatus icarus (Lepidoptera: Lycaenidae). Canadian Journal of Zoologypdf   press release  CBC Ottawa Morning interview   Le Devoir article

Lalonde, S., McCune, J.L., Rivest, S.A., and Kharouba, H.M. 2021. Decline in common milkweed along roadsides around Ottawa, Canada. Écosciencepdf

Tremblay, P.T., MacMillan, H.A., and Kharouba, H.M. 2021. Autumn larval cold tolerance does not predict the northern range limit of a widespread butterfly species. Ecology and Evolution. pdf

Dargent, F., Gilmour, S.M., Brown, E.A., Kassen, R. and Kharouba, H.M. 2021. Low prevalence of the parasite Ophryocystis elektroscirrha at the range edge of the eastern North American monarch (Danaus plexippus) butterfly population. Canadian Journal of Zoology.  pdf

2015-2020

Kharouba, H.M. and Wolkovich, E.M. 2020. Disconnects between ecological theory and data in phenological mismatch research. Nature Climate Change, 10:406-415. doi: 10.1038/s41558-020-0752-x. link pdf Nature Ecology & Evolution blog post

McCune, J.L., Roser-Katz, H., Bennett, J.R., Schuster, R., & Kharouba, H.M. 2020. Do traits of plant species predict the efficacy of species distribution models for finding new occurrences? Ecology and Evolution, 00:1-14. DOI:10.1002/ece3.6254. pdf

Radchuk, V. + 64 co-authors (incl. Kharouba, H.M). 2019. Adaptive responses of animals to climate change are most likely insufficient. Nature Communications 10: 3109. pdf

Panchen, Z.A., Doubt, J.M, Kharouba, H.M., and Johnston, M.O. 2019. Patterns and biases in an Arctic herbarium specimen collection: implications for phenological research. Applications in Plant Sciences 7:e1229. Special Issue: “Emerging Frontiers in Plant Phenology”.  pdf

Chmura, H.E., Kharouba, H.M., Ashander, J., Ehlman, S.M., Rivest, E.B., and Yang, L.H. 2019. The mechanisms of phenology: the patterns and processes of phenological shifts. Ecological Monographs 89:e01337. DOI: 10.1002/ecm.1337. pdf

​Kharouba, H.M., Lewthwaite, J.M.M., Guralnick, R., Kerr, J.T., and Vellend, M. 2018. Using insect natural history collections to study global change impacts: challenges and opportunities. Theme Issue: Biological collections for understanding biodiversity in the Anthropocene. Philosophical Transactions B 374: 20170405. pdf

Kharouba, H.M., Ehrlen, J., Gelman, A., Bolmgren, K., Allen, J., Travers, S. and E.M. Wolkovich. 2018. Global shifts in the phenological synchrony of species interactions over recent decades. PNAS  115:5211-5216.  pdf  Commentary; Media coverage: National PostCBC Radio-CanadaNew York Times

Buckley, L.B., Arakaki, A.J., Cannistra, A.F., Kharouba, AH.M. and Kingsolver, J.G. 2017. Insect development, thermal plasticity and fitness implications in changing, seasonal environments. Integrative and Comparative Biology, icx032.  pdf

Wetzel, W.C., Kharouba, H.M., Robinson, M., Holyoak, M. and Karban, R. 2016. Variance in plant nutritive traits reduces the performance of insect herbivores. Nature 539:425.  link  data

Lee-Yaw, J.A.,  Kharouba, H.M, Bontrager, M., Mahony, C.,  Csergő, A.M,  Noreen, A.M.E., Li, Q., Schuster, R., and Angert, A.L. 2016.  A synthesis of transplant experiments and ecological niche models suggests that range limits are often niche limits. Ecology Letters 19:710-722. pdf

2010-2015

Karban, R., Grof-Tisza, P., McMunn, M., Kharouba, H.M., and Huntzinger, M. 2015. Caterpillars escape predation in habitat and thermal refuges.  Ecological Entomology 40:725-731.  pdf

Kharouba, H.M. and Vellend M. 2015. Flowering time of butterfly nectar food plants is more sensitive to temperature than the timing of butterfly adult flight. Journal of Animal Ecology 84:1311-1321.  pdf

Kharouba, H.M., Vellend, M., Sarfraz, R. and Myers, J.H. 2015. The effects of experimental warming on the timing of a plant-insect herbivore interaction. Journal of Animal Ecology 84:785-796.  pdf

Gilbert, B., Tunney, T.D., McCann, K.S., DeLong, J.P. Vasseur, D.A., Savage, V., Shurin, J.B., Dell, A.I., Barton, B.T., Harley, C.D.G., Kharouba, H.M., Kratina, P., Blanchard, J.L., Clements, C., Winder, M., Greig, H.S. and O’Connor, M.I. 2014. A bioenergetic framework for the temperature dependence of trophic interactions. Ecology Letters 17:891-1052. pdf

Kharouba, H.M., Paquette, S.R., Kerr, J.T and Vellend, M. 2014. Predicting the sensitivity of butterfly phenology to temperature over the past century. Global Change Biology 20:504-514.  pdf; press coverage: press releaseDaily Planet (Discovery Channel Canada), Vancouver 24hrsthe Conversation (uk); related stories: cbc (the National and article)

Vellend, M. and Kharouba, H.M. 2013. Setting conservation priorities when what you see is not what you get. Animal Conservation 16:14-15. (Invited commentary).  pdf

Vellend, M., Brown, C., Kharouba, H.M., McCune, J., and Myers-Smith, I. 2013. Historical ecology: using unconventional data sources to test for impacts of global environmental change. American Journal of Botany 100:1294-1305pdf

Sarfraz, R., Kharouba, H.M., and Myers, J.H. 2013. Tent caterpillars are robust to variation in leaf phenology and quality in two thermal environments. Bulletin of Entomological Research 103:522-529.  pdf

Kharouba, H.M., McCune, J.L., Thuiller, W., and Huntley, B. 2013. Do ecological differences between taxonomic groups influence the relationship between species’ distributions and climate? A global meta-analysis using species distribution models. Ecography 36:657-664. Editor’s Choice.  pdf 

Williams, J.W., Kharouba, H.M., Veloz, S., Vellend, M., McLachlan, J.,  Liu, Z., Otto-Bliesner, B., and He, F. 2013. The Ice Age Ecologist: Testing Methods for Reserve Prioritization During the Last Global Warming. Global Ecology and Biogeography 22:289-301. Faculty of 1000 “Recommended”. pdf   review

Kharouba, H.M., and Kerr, J.T. 2010. Just passing through: Global change and the conservation of biodiversity in protected areas. Biological Conservation 143:1094-1011. pdf

Kharouba, H.M., Algar, A.C. and Kerr, J.T. 2009. Historically calibrated predictions of butterfly species’ range shift using global change as a pseudo-experiment. Ecology 90:2213-2222. pdf

Algar, A.C., Kharouba, H.M., Young, E.R., and Kerr, J.T. 2009. Predicting the future of species diversity: macroecological theory, climate change, and direct tests of alternate forecasting methods. Ecography 32:22-33. pdf

Kharouba, H.M., Nadeau, J.L., Young, E., and Kerr, J.T. 2008. Using species distribution models to effectively conserve biodiversity into the future. Biodiversity, Special Issue: Global change and biodiversity 9:39-46. pdf

Kerr, J.T., Kharouba, H.M., and Currie, D.J. 2007. The macroecological contribution to global change solutions. Science 316:1581-1584. pdf

Book chapters
Kerr, J.T. and Kharouba, H.M. 2007. Climate change and conservation biology. In: May, R.M. and Maclean, A. (eds). Theoretical Ecology. 3rd edition. Oxford University Press, Oxford, pp 190-204.

2007 - 2010